Basically it'd be a kind of circle of life I guess.
Essentially, all organic enemies will try and kill each other on sight (to a degree)
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Brackens are neutral
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Hoardbugs attack any enemy near their nest or that attacks them
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Spiders kill anything that hits their web (or try at least)
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Slimes kill anything except Brackens, imagine the Brackens are vaulting over the slime
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Thumpers kill on sight but flee from slimes. Can be lured to charge into slimes killing them
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Coil heads don't care about anything else since they are concrete and focus on you only
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Jesters kill anything in their path towards you
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Ghost Girl doesn't do anything because she isn't real, and cannot hurt you.
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Eyeless dogs will try and kill giants (failing of course)
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Worms will kill the closest thing to them, giant or otherwise.
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Bees kill anything near their hive except worms and giants